Agalenocosa Mello-Leitão, 1944

"Medium-sized wolf spiders (TL, 4.20–9.98 mm). Females larger than males. Carapace brown with a light brown median band and a Y-shaped mark that extends from fovea to the PLE (tenuous in A. grismadoi sp. nov. and A. punctata, absent in A. luteonigra). Eyes surrounded by black pigment. Dorsum of abdomen brown to dark olive-grey, with light yellow lanceolate mark in the cardiac area and a pattern of white spots laterally and posterior to the cardiac mark, composed by glistening setae (Fig. 3b) (except in A. luteonigra, which have a black
abdomen). Carapace longer than wide, dorsal line straight in lateral view (Figs 9b, e; domed in A.grismadoi sp. nov. – Figs 18b, e). Caput flanks in frontal view steep in males, gently sloping in females. AME larger than ALE, AER slightly procurved in anterior view (Fig. 24). Chelicerae with three promarginal and three retromarginal teeth. Abdomen oval. Colulus a fleshy triangular lobe, with several setae (fig. 5a). The spinnerets have a similar conformation to what Townley and Tillinghast (2003) reported for Lycosidae, with six spinnerets, ALS and PLS two segmented, PMS with a single segment (Figs 5a, 6a). ALS with two major ampullate gland spigots on the mesal margin (the posterior one reduced to a nubbin in the male (Fig. 6b)) and piriform gland spigots (Fig. 5b). PMS with about 30 aciniform and two cylindrical gland spigots in the female (Fig. 5c) and about 20 aciniform in the male and without cylindrical gland spigots (Fig. 6c), with a few setae between them. Female with two minor ampullate gland spigot with a nubbin and a tartipore close to them on the PMS (Fig. 5c) and only one minor ampullate gland spigot in the male (Fig. 6c). PLS long, tubular, distal segment short, conical, with aciniform and cylindrical gland spigots, with setae among them (Fig. 5d). Leg formula IV > I > II > III or IV > I > III > II, except in A. luteonigra where the leg formula is IV >III > I >
II. Tibia I with two pairs of ventral spines (2-2-0), except in A. pirity sp. nov. (0-2-0) and A. grismadoi sp. nov. (2-0-0). Femur of male palp without modifications. Tibia with a basal retrolateral apophysis, covered with strong setae (Fig. 3a), reduced in A. grismadoi sp. nov. (Fig. 8f). Cymbium symmetric, without macrosetae in its tip. Tegulum with a prolateral furrow, deep in A. velox Keyserling, 1891 (Figs 2a, 10d) and A. tricuspidata Tullgren, 1905 (Fig. 12d) and shallow in the rest of the species. Median apophysis membranous, composed of two elements, the ventral one is flattened and directed longitudinally, the dorsal is transverse and usually retrolaterally directed (Fig. 2a, b). The embolic division possesses a grooved and complex terminal apophysis, with a pointed tip (Figs 2a, b, 7) and two additional processes, one ventral and a dorsal (Fig. 2). Embolus as a curved spine, not visible in the unexpanded palp, originating prolaterally, below the ventral process of the embolic division and running retrolaterally hidden by the terminal apophysis (Fig. 7). Epigyne composed of a simple plate with two soft sclerotised areas; in A. velox, A. punctata, A. gentilis and A. grismadoi sp. nov. this area is well developed and surrounds the copulatory open, and in A. luteonigra, A. pirity sp nov., A. tricuspidata and A. gamas sp. nov. is reduced to a small area, posterior to the copulatory open. Vulva: spermathecae with a distal head connected to the base by a more or less defined stalk, vulval chambers rounded. Dictynoid pore conspicuous, located on the boundaries of the vulval chamber and the spermathecal stalk (Fig. 4a)." Piacentini, 2014